phlb = parasitoid host location behaviour
This read me file was made for "experiment 2" as described in: 

Croijmans, L., Valstar, R. T., Schuur, L., Jacobs, I., van Apeldoorn, D. F., & Poelman, E. H. (2022). 
Intraspecific plant variation and nonhost herbivores affect parasitoid host location behaviour. Animal Behaviour, 194, 169-184.

For a full explanation of the methods, see above paper.

EXPERIMENTAL DESIGN
In this experiment, we examined parasitoid foraging behaviour within a wind-tunnel (200 x 60 x 60 cm) 
to assess how intraspecific plant variation and host presence affect foraging efficiency of the parasitoid Cotesia glomerata to its host Pieris brassicae.
Plant rows differed in the composition of cultivars (more attractive Christmas Drumhead (C) and less attractive Rivera (R) cultivars) and host presence (absence, only present at last plant, present at all plant).

We tested 10 different plant / host combinations:
1. RRRR			--> All plants are Rivera without herbivores
2. RRRC			--> First three plants are Rivera, last plant Christmas Drumhead. All withouth herbivores.
3. RRRRH		--> All plants are Rivera, only last plant has herbivores.
4. RRRCH		--> First three plants are Rivera, last plant Christmas Drumhead. Only last plant has herbivores.
5. CCCRH		--> First three plants are Christmas Drumhead, last plant Rivera. Only last plant has herbivores.
6. CCCCH		--> All plants are Christmas Drumhead, only last plant has herbivores.
7. RHRHRHRH		--> All plants are Rivera with herbivores.
8. RHRHRHCH		--> First three plants are Rivera, last plant Christmas Drumhead. All with herbivores.
9. CHCHCHRH		--> First three plants are Christmas Drumhead, last plant Rivera. All with herbivores.
10.CHCHCHCH		--> All plants are Christmas Drumhead with herbivores.


MEASUREMENTS
We used a hand-held computer with the ‘The Observer XT 10.1’ software (Noldus Information Technology) to track parasitoid behaviour. 
We recorded five behavioural parameters: the proportion of parasitoids that initiated flight and landed in the plant row (Response); 
the proportion of parasitoids that flew directly to the last plant in a row (Direct Flight); 
the time the parasitoid took to reach the last plant from the first time it took flight (Searching Time); 
the time the parasitoid spent on the first three plants in a row (Plant 123 Time); 
and the time it took the parasitoid to attack the hosts after landing on the last plant for the first time (Attack Latency).
The time the parasitoid spent on each plant was tracked separately until the parasitoid successfully parasitized a larva on the last plant 
(if no larvae were present at all, recording was stopped once the parasitoid reached the last plant), or until 30 min had elapsed. 
Any parasitoids that were responsive but did not reach the last plant within 30 min were considered to have taken the full 30 min to get to the last plant, 
thus underestimating time to reach the last plant. If a parasitoid remained motionless on a plant for more than 5 min, it was recorded as an ‘unsuccessful forager’ and the recording was terminated. 
If the parasitoid stayed in the glass cylinder or on one of the sides of the wind tunnel for more than 5 min without touching one of the plants, 
the recording was terminated, and the parasitoid was considered ‘unresponsive’.


RAW DATA
Data in csv file is semicolon delimited.

Colums have the following definition:

Observation file			= Unique ID number for observation.
Total duration in glass cylinder	= Number of seconds the parastioid spend in the glass cylinder that served as starting point (after around 300 seconds, measurements were terminated).
Total duration flight			= Number of seconds the parastioid spend flying.
Total number of flights			= Number of times the parasitoid was flying in between landings.
Total duration stay on plant 1/2/3/4	= Number of seconds the parastioid spend sitting/walking on respectively plant 1, 2, 3 or 4.
% duration stay on plant 1/2/3/4	= Percentage of time spend on plants, that the parasitoid spend on respectively plant 1, 2, 3 or 4.
Total number of stays on plant 1/2/3/4	= Number of times the parasitoid landed on respectively plant 1, 2, 3 or 4.
Total duration stay on wind tunnel	= Number of seconds the parastioid spend sitting/walking on the walls, roof or floor of the wind tunnel.
Total number of stays on wind tunnel	= Number of times the parasitoid landed on the walls, roof or floor of the wind tunnel. 
Total duration of stays on a plant	= Total number of seconds the parasitoid spend on all plants.
Total duration of stays			= Total number of seconds the parasitoid spend on plants and on the walls, roof or floor of the wind tunnel.
Total number of stays on plants		= Total number of times the parasitoid landed on any plant.
Total number of stays			= Total number of times the parasitoid landed on any plant or the walls, roof or floor of the wind tunnel.
Total number of attacks on plant 1/2/3/4= Total number of times the parasitoid attacked caterpillars on respectively plant 1, 2, 3 or 4. 
						Observation was terminated after attack the parasitoid attacked a caterpillar on plant 4 (or when it reached plant 4 in the case of RRRR and RRRC).
						Attacks on caterpillars on plants 1, 2 and 3 was only possible for treatments 7 to 10. 
Total number of attacks			= Total number of times the parasitoid attacked caterpillars on any plant.
First three plants			= Cultivar identity (R = Rivera, CD = Christmas Drumhead) and herbivore presence (clean = absent, induced = present) of the first three plants
Last plant				= Cultivar identity (R = Rivera, CD = Christmas Drumhead) and herbivore presence (clean = absent, induced = present) of the last plant.
Name Observer				= Name of the observer that did the measurement
Total duration				= Total number of seconds until observation was terminated.
Start time				= Time of the day that the observation was started.
Stop time				= Time of the day that the observation was terminated.
Response				= Binary variable, indicating whether the parasitoid landed on any plant (1 = it did, 0 = it did not).
Flying					= Binary variable, indicating whether the parasitoid initiated flight (1 = it did, 0 = it did not).
Succesfull				= Binary variable, indicating whether the parasitoid succesfully parasitized a caterpillar on (or reached in the case of RRRR and RRRC) plant 4. 
First plant landing			= Nominal variable, indicating which plant the parasitoid landed first on.
Landing on plant 4			= Binary variable, indicating whether the parasitoid landed on plant 4 (1 = it did, 0 = it did not).
Time until reach plant 4		= Number of seconds it took the parasitoid to land on plant 4 for the first time, since being released.
Time until attacking on plant 4		= Number of seconds it took the parasitoid to parasitize a caterpillar on plant 4 for the first time.
DirectFlight				= Binary variable, indicating whether the parasitoid landed first on plant 4 out of all other plants (1 = it did, 0 = it did not).
Treatment				= Treatments as mentioned above.
TrCode					= Treatment codes as mentioned above.
Cultivar123				= Binary variable, indicating the identity of the first three plants (0 = Rivera, 1 = Christmas Drumhead).
Host123					= Binary variable, indicating the presence of herbivores on the first three plants (0 = absent, 1 = present).
Cultivar4				= Binary variable, indicating the identity of the last plant (0 = Rivera, 1 = Christmas Drumhead).
Host4					= Binary variable, indicating the presence of herbivores on the last plant (0 = absent, 1 = present).
Date					= Date the observation was on, in the DD-MM-YYYY format.